Curriculum Vitae
JONATHAN MARK ADRAIN
Business Address: Department of Geoscience
University of Iowa, Iowa City, Iowa 52242
Phone: 319-335-1539
E-mail: jonathan-adrain@uiowa.edu
EDUCATIONAL AND PROFESSIONAL HISTORY
1. Higher Education
1993, PhD, Geology, University of Alberta, Edmonton, Alberta
1989, BSc, Geology and Zoology, University of Alberta, Edmonton, Alberta
2. Professional and Academic Positions
2004-present, Associate Professor, Department of Geoscience, University of Iowa
1999-2004, Assistant Professor, Department of Geoscience, University of Iowa
1998-1999, Postdoctoral Research Assistant, Leverhulme Trust Grant, Department of Palaeontology, Natural History Museum, London
1995-1998, Postdoctoral Research Assistant, Natural Environment Research Council (UK) Grant, Department of Palaeontology, Natural History Museum, London
1993-1995, Postdoctoral Fellow, Natural Sciences and Engineering Research Council (Canada), Department of Earth Sciences, University of Western Ontario
3. Honors and Awards
2004 Pikaia Award, Geological Association of Canada, Paleontology Division
2004 Dean's Scholar, College of Liberal Arts and Sciences, Class of 2004-2006
4.
Memberships
Palaeontological Association
Paleontological Society
Society of Systematic Biologists
Willi Hennig Society
SCHOLARSHIP
1. Publications (*=major contribution; **=secondary
contribution; ***=equal major
contribution; ****=minor contribution)
a. Refereed
Ą Monographs
(3) Jell, P. A., and Adrain ***, J. M. 2003. Available generic names for trilobites. Memoirs of the Queensland Museum, 48:331-558.
(2) Adrain, J. M. 1997. Proetid trilobites from the Silurian (Wenlock-Ludlow) of the Cape Phillips Formation, Canadian Arctic Archipelago. Palaeontographia Italica, 84:1-91.
(1) Adrain*, J. M., and Edgecombe, G. D. 1997. Silurian encrinurine trilobites from Arctic Canada. Palaeontographica Canadiana, 14:1-109.
Ą Edited
Volumes
(2) Adrain, J. M., Edgecombe, G. D., and Lieberman, B. S. (eds). 2001. Fossils, Phylogeny, and Form: An Analytical Approach. Kluwer Academic/Plenum Press, New York, 402p.
(1) Adrain, J. M., and Westrop, S. R. (eds). 1999. Trilobite Paleobiology. [Special theme issue of the Journal of Paleontology, comprising selected papers from the Second International Trilobite Conference.] Journal of Paleontology, 73(2):161-371.
Ą
Articles
(46) Adrain, J. M.* and Westrop, S. R. 2007. Notchpeakia, a new genus of Upper Cambrian (Sunwaptan) "entomaspidid" trilobites. Journal of Paleontology, 81, in press.
(45) Adrain, J. M.* and Westrop, S. R. 2006. New genus of aulacopleuroidean trilobites from the earliest Ordovician of Laurentia. Acta Palaeontologica Polonica, 51, in press.
(44) Adrain, J. M.* and Westrop, S. R. 2006. New earliest Ordovician trilobite genus Millardicurus: the oldest known hystricurid. Journal of Paleontology, 80, in press.
(43) Adrain, J. M.* and Tetreault, D. K. 2005. The brachymetopid trilobite Radnoria in the Silurian (Wenlock) of New York State and Arctic Canada. Canadian Journal of Earth Sciences, 42, in press.
(42) Adrain, J. M.*. and Westrop, S. R. 2005. Lower Ordovician trilobites from the Baumann Fiord Formation, Ellesmere Island, Arctic Canada. Canadian Journal of Earth Sciences, 42, in press.
(41) Adrain, J. M.***, and Westrop, S. R. 2005. Ptychaspidid trilobites from the Upper Cambrian (Sunwaptan) Notch Peak Formation, Utah. Geological Magazine, 142, in press.
(40) Adrain, J. M. 2005. Aulacopleurid trilobites from the Upper Ordovician of Virginia. Journal of Paleontology, 79:548-569.
(39) Adrain, J. M.*, Edgecombe, G. D., Fortey, R. A., Laurie, J, McCormick, T., Owen, A. W., Waisfeld, B, Webby, B. D., Westrop, S. R. & Zhou, Z.-Y. 2004. Trilobites. In Webby, B. D., Droser, M. L., and Paris, F. (eds.), The Great Ordovician Biodiversification Event. Columbia University Press, New York.
(38) Adrain, J. M. 2003. Validity and composition of the Silurian trilobite genera Borealarges and Dicranogmus, with new species from the Canadian Arctic. Canadian Journal of Earth Sciences, 40:749-763.
(37) Adrain, J. M.*** & Westrop, S. R. 2003. Paleobiodiversity: We need new data. Paleobiology, 29:22-25.
(36) Adrain*, J. M., Lee, D.-C., Westrop, S. R., Chatterton, B. D. E., and Landing, E. 2003. Classification of the trilobite subfamilies Hystricurinae and Hintzecurinae subfam. nov., with new genera from the Lower Ordovician (Ibexian) of Idaho and Utah. Memoirs of the Queensland Museum, 48:559-592.
(35) Westrop, S. R., and Adrain**, J. M. 2001. Sampling at the species level: impact of spatial biases on diversity gradients. Geology, 29:903-906.
(34) Adrain*, J. M., Edgecombe, G. D., and Lieberman, B. S. 2001. Paleobiology: An empirical historical science. Pp. 1-6 in Adrain, J. M., Edgecombe, G. D., and Lieberman, B. S. (eds), "Fossils, Phylogeny, and Form: An Analytical Approach." Kluwer Academic/Plenum Press, New York.
(33) Adrain***, J. M., and Westrop, S. R. 2001. Stratigraphy, phylogeny, and species sampling in time and space. Pp. 291-322 in Adrain, J. M., Edgecombe, G. D., and Lieberman, B. S. (eds), "Fossils, Phylogeny, and Form: An Analytical Approach." Kluwer Academic/Plenum Press, New York.
(32) Brochu, C. A., Bryant, H. N., Theodor, J. M., O'Leary, M. A., Adrain, J. M.****, and Sumrall, C. D. 2001. Modern phylogenetics in paleontology: comments on Vermeij. Paleobiology, 27:174-176.
(31) Adrain*, J. M., Westrop, S. R., Landing, E., and Fortey, R. A. 2001. Systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella, with species from the type Ibexian area, western U.S.A. Journal of Paleontology, 75:947-971.
(30) Adrain*, J. M., Westrop, S. R. 2000. An empirical assessment of taxic paleobiology. Science, 289:110-112.
(29) Adrain***, J. M., Westrop, S. R., Chatterton, B. D. E., and Ramskšld, L. 2000. Silurian trilobite alpha diversity and the end-Ordovician mass extinction. Paleobiology, 26: 625-646.
(28) Hughes, N. C., Chapman, R. E., and Adrain****, J. M. 1999. The stability of thoracic segmentation in trilobites: A case study in developmental and ecological constraints. Evolution and Development, 1:24-35.
(27) Adrain*, J. M., Fortey, R. A., and Westrop, S. R. 1998. Post-Cambrian trilobite diversity and evolutionary faunas. Science, 280:1922-1925.
(26) Adrain, J. M. 1998. Systematics of the Acanthoparyphinae (Trilobita), with species from the Silurian of Arctic Canada. Journal of Paleontology, 72:698-718.
(25) Westrop, S. R., and Adrain**, J. M. 1998. Trilobite diversity and the Ordovician Radiation: a reply. Paleobiology, 24:529-533.
(24) Westrop, S. R., and Adrain***, J. M. 1998. Trilobite alpha diversity and the reorganization of Ordovician benthic marine communities. Paleobiology, 24:1-16.
(23) Davidek, K., Landing, E., Bowring, S. A., Westrop, S. R., Rushton, A. W. A., Fortey, R. A., and Adrain****, J. M. 1998. New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian-Ordovician boundary. Geological Magazine, 135:305-309.
(22) Adrain*, J. M., and Fortey, R. A.. 1997. Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum (Geology), 53:81-117.
(21) Adrain*, J. M., and Kloc, G. J. 1997. Lower Devonian aulacopleuroidean trilobites from Oklahoma. Journal of Paleontology, 71:703-712.
(20) Adrain***, J. M., and Edgecombe, G. D. 1997. Silurian calymenid trilobites from the Cape Phillips Formation, central Canadian Arctic. Journal of Paleontology, 71:657-682.
(19) Adrain*, J. M., and Ramskšld, L. 1997. Silurian odontopleurine trilobites from the Cape Phillips Formation, central Canadian Arctic. Journal of Paleontology, 71:237-261.
(18) Adrain*, J. M., and MacDonald, E. W. 1996. Phacopid trilobites from the Silurian of Arctic Canada. Journal of Paleontology, 70:1091-1094.
(17) Adrain, J. M. 1996. A new otarionine trilobite from the Henryhouse Formation (Silurian, Ludlow) of Oklahoma. Journal of Paleontology, 70:611-614.
(16) Adrain*, J. M., and Ramskšld, L. 1996. The lichid trilobite Radiolichas in the Silurian of Arctic Canada and Gotland, Sweden. Geological Magazine, 133:147-158.
(15) Adrain*, J. M., and Edgecombe, G. D. 1996. Devonian aulacopleurid trilobites of the Malvinokaffric Realm. Geobios, 29:417-436.
(14) Adrain*, J. M., and Chatterton, B. D. E. 1996. The otarionine trilobite Cyphaspis, with new species from northwestern Canada. Journal of Paleontology, 70:100-110.
(13) Adrain*, J. M., Chatterton, B. D. E., and Blodgett, R. B. 1995. Silurian trilobites from southwestern Alaska. Journal of Paleontology, 69:723-736.
(12) Adrain***, J. M., and Edgecombe, G. D. 1995. Balizoma and the new genera Aegrotocatellus and Perirehaedulus: encrinurid trilobites from the Douro Formation (Silurian, Ludlow) of the central Canadian Arctic. Journal of Paleontology, 69:736-752.
(11) Adrain*, J. M., and Chatterton, B. D. E. 1995. Aulacopleurine trilobites from the Llandovery of northwestern Canada. Journal of Paleontology, 69:326-340.
(10) Adrain*, J. M., and Chatterton, B. D. E. 1995. The otarionine trilobites Harpidella and Maurotarion, with species from northwestern Canada, the United States, and Australia. Journal of Paleontology, 69:307-326.
(9) Edgecombe, G. D., and Adrain****, J. M. 1995. Silurian calymenid trilobites from the United States. Palaeontographica Abt. A, 235:1-19.
(8) Adrain*, J. M., and Wilson, M. V. H. 1994. Early Devonian cephalaspids (Vertebrata: Osteostraci: Cornuata) from the southern Mackenzie Mountains, N.W.T., Canada. Journal of Vertebrate Paleontology, 14:301-319.
(7) Adrain, J. M. 1994. The lichid trilobite Borealarges n. gen., with species from Arctic Canada. Journal of Paleontology, 68:1081-1099.
(6) Adrain*, J. M., and Chatterton, B. D. E. 1994. The aulacopleurid trilobite Otarion, with new species from northwestern Canada. Journal of Paleontology, 68:305-323.
(5) Ramskšld, L., Adrain***, J. M., Edgecombe, G. D., and Siveter, D. J. 1994. The calymenid trilobite Alcymene n. gen., with species from the Ludlow of Gotland, Sweden. Journal of Paleontology, 68:556-569.
(4) Adrain*, J. M., and Chatterton, B. D. E. 1993. A new rorringtoniid trilobite from the Ludlow of Arctic Canada. Canadian Journal of Earth Sciences, 30:1634-1643.
(3) Adrain, J. M. 1992. Machaeridian classification. Alcheringa, 16:15-32.
(2) Adrain*, J. M., Chatterton, B. D. E., and Cocks, L. R. M. 1991. A new species of machaeridian from the Silurian of Podolia, USSR, with a review of the Turrilepadidae. Palaeontology, 34:637-651.
(1) Adrain*, J. M., and Chatterton, B. D. E. 1990. Odontopleura (Trilobita, Silurian), and a method of constrained congruency analysis. Journal of Paleontology, 64:600-614.
b. Non-refereed
(6) Adrain, J. M. 2004. The Cherry Creek Trilobites. American Paleontologist 12(4):6-9.
(5) Adrain, J. M. 2001. Systematic paleontology. Journal of Paleontology, 75:1055-1057.
(4) Adrain, J. M. and Edgecombe, G. D. 2001. Trilobites. McGraw-Hill Encyclopedia of Science and Technology, 9th Edition. (invited contribution)
(3) Adrain, J. M., and Westrop, S. R. 1999. Trilobite paleobiology: Past, present, and future. Journal of Paleontology, 73:161-162.
(2) Adrain, J. M. 1999. (book review) "Arthopod Fossils and Phylogeny." Columbia University Press. Paleontologica Electronica 2(1).
(1) Adrain, J. M. 1997. (book review) "Trilobite systematics, biostratigraphy, and palaeoecology of the Lower Ordovician Komstad Limestone and Huk Formations, southern Scandinavia." Palaeontology Newsletter, 30:26-27.
3. Research
Grants
a. External
As PI:
2003-2006, National Science Foundation Grant EAR-0308685, 8/01/2003-7/31/2006, $115,000; "Collaborative Research: Patterns of diversity and faunal turnover at a Late Cambrian trilobite extinction event: testing for phylogenetic, stratigraphic and environmental biases". (PI=J.M. Adrain)
As
Co-PI:
2001-2003, National Science Foundation Grant DBI-0096768 (2 yrs., 10/1/01-9/30/03, $255,149; "Reorganization of the University of Iowa Paleontology Repository". (PI=A. F. Budd; co-PIs = J. M. Adrain, J. Golden)
1999-2002 Subcontract on National Science Foundation Grant EAR-9973065 (3 yrs., 8/1/99-7/31/02, full grant amout $179,999; Iowa subcontract amount = $72,000); (PI = Steve Westrop, University of Oklahoma; co-PI = J. M. Adrain)
b. Internal
2003-2004 Iowa Research Experiences for Undergraduates grant
2002 Old Gold Summer Fellowship, University of Iowa
2000 Old Gold Summer Fellowship, University of Iowa
2000 Carver Scientific Research Initiative, University of Iowa, $12,572
5. Conference Presentations (+=invited, * = speaker, P=poster)
a. International
(8) Adrain, J. M. 2004. Taphonomic control of shallow subtidal fossil abundance. Fourteenth Canadian Paleontology Conference (CPC 2004). St. Andrews, New Brunswick, Canada, September 2004.
(7) Adrain, J. M. 2003. Silicified trilobites from the Upper Cambrian (Sunwaptan) Notch Peak Formation, Utah: Implications for Cambrian trilobte systematics and biostratigraphy. Thirteenth Canadian Paleontology Conference (CPC-2003). Edmonton, Alberta, Canada, September 2003.
(6+) Adrain, J. M. 2002. Time and phylogeny reconstruction. 2002 Lyell Meeting: Methods of Phylogenetic Reconstruction. London, England, June 2002.
(5) Adrain, J. M.*, and Westrop, S. R. 2001. Trilobite alpha diversity: decoupling taxonomic and ecologic "success". Third International Conference on Trilobites and their Relatives, Oxford, England, April 2001.
(4+) Adrain, J. M.*, and Westrop, S. R. 2001. The "cryptogenesis" problem and higher trilobite phylogeny. Third International Conference on Trilobites and their Relatives, Oxford, England, April 2001.
(3) Miller, G. C., and Adrain, J. M. 1997. Conodonts from the Cape Phillips Formation (Wenlock, Silurian) of Arctic Canada. Ichthyolith Issues Special Publication No. 3: Palaeozoic Strata and Fossils of the Eurasian Arctic, p. 29-30.
(2) Adrain, J. M.* 1996. Trilobites, phylogenetic context, and the Palaeozoic Evolutionary Fauna. Palaeontological Association Annual Conference, Birmingham, England. Palaeontology Newsletter, 32:10.
(1) Adrain, J. M.*, and Edgecombe, G. D. 1995. Silurian calymenid trilobites from eastern and Arctic North America. Palaeontological Association Annual Conference, Galway, Ireland. Palaeontology Newsletter, 28:9.
b. National
(26) Adrain, J. M.*, and Westrop, S. R. 2005. Evenness and diversity of Lower Paleozoic trilobite faunas of Laurentian North America. Geological Society of America Abstracts with Programs, 36(5):455.
(25P) Adrain, T. S., Budd, A. F., Adrain, J. M., and Golden, J. 2005. UI Paleontology Repository reorganization project: Improving standards in collection care and access. Geological Society of America Abstracts with Programs, 36(5):455.
(24) Adrain, J. M.*, and Westrop, S. R. 2004. Taphonomic control of shallow subtidal fossil abundance. Geological Society of America Abstracts with Programs, 37.
(23) Adrain, J. M., and Westrop, S. R.* 2004. Rapid turnover of trilobite faunas related to stratigraphic condensation in a Lower Ordovician "biomere". Geological Society of America Abstracts with Programs, 36(5):526.
(22) Hegna, T.*, and Adrain, J. M. 2004. Reduced biofacies differentiation in Silurian trilobite faunas of Laurentia. Geological Society of America Abstracts with Programs, 36(5):177.
(21P) Ng, T.-W., and Adrain, J. M. 2004. Upper Marjuman (Upper Cambrian) shallow water trilobite faunas, Great Basin, western USA. Geological Society of America Abstracts with Programs, 36(5):364.
(20P) Karim, T., and Adrain, J. M. 2004. Lower Ordovician trilobites of the Cow Head Group, western Newfoundland, Canada. Geological Society of America Abstracts with Programs, 36(5):365.
(19P) Hegna, T., and Adrain, J. M. 2003. Early Silurian trilobites from northeastern Missouri. Geological Society of America Abstracts with Programs.
(18) Adrain, J. M.*, and Westrop, S. R. 2001. Early Ordovician trilobite diversity, turnover rates, and mass extinction: High-resolution biostratigraphy of the House Formation and correlatives, Great Basin. Geological Society of America Abstracts with Programs, 33(6).
(17) Westrop, S. R., Adrain, J. M., and Bean, J. R. 2001. Stratigraphic context of Late Cambrian trilobite extinctions: Evidence from eastern Nevada. Geological Society of America Abstracts with Programs, 33(6).
(16P) Schultz, D. B., Adrain, J. M., and Westrop, S. R. 2001. Biostratigraphy and paleoecology of Late Cambrian (Marjuman) trilobites from the Emigrant Springs Formation, southern Schell Creek Range, Nevada. Geological Society of America Abstracts with Programs, 33(4).
(15) Bean, J., Adrain, J. M., and Westrop, S. R. 2001. Trilobite paleoecology and taphonomy of the Steptoean (Upper Cambrian) Dunderberg Formation, Cherry Creek Range, Nevada. Geological Society of America Abstracts with Programs, 33(4).
(14) Adrain, J. M.* 2000. Regional Silurian trilobite turnover coincident with graptolite mass extinction. Geological Society of America Abstracts with Programs, 32(7):A367.
(13) Westrop, S. R., and Adrain, J. M. 2000. Sampling at the species level: Impact of spatial and temporal biases on diversity gradients. Geological Society of America Abstracts with Programs, 32(7):A131.
(12) Adrain, J. M.*, and Westrop, S. R. 1999. Ontogeny, cryptogenesis, and the origin of the Whiterock trilobite fauna. Geological Society of America Abstracts with Programs, 31(7):A138.
(11) Hughes, N. C., Chapman, R. E., and Adrain, J. M. 1999. Developmental and ecological constraints in the aftermath of Cambrian Radiation: The trilobite thorax as a test case. Geological Society of America Abstracts with Programs, 31(7):A363.
(10) Adrain, J. M.*, Fortey, R. A., and Westrop, S. R. 1998. The Ordovician radiation of trilobites. Geological Society of America Abstracts with Programs, 30(7):A285.
(9) Adrain, J. M.*, Westrop, S. R., Chatterton, B. D. E., and Ramskšld, L. 1998. Silurian trilobite alpha diversity and the end-Ordovician extinction. Geological Society of America Abstracts with Programs, 30(7):A285.
(8) Adrain, J. M.*, Fortey, R. A., and Westrop, S. R. 1997. Trilobite diversity patterns: implications for paleobiological analysis. Geological Society of America Abstracts with Programs, 29(7).
(7) Adrain, J. M.* 1996. Measures of diversity, the Paleozoic Evolutionary Fauna, and the reinvention of trilobites. Geological Society of America Abstracts with Programs, 28(7):A291.
(6) Adrain, J. M.*, and Fortey, R. A. 1996. The basal Middle Ordovician faunal revolution in Laurentia: phylogeny and paleoenvironment. Sixth North American Paleontological Convention Abstracts of Papers. Paleontological Society Special Publication, 8:1.
(5) Adrain, J. M.*, and Westrop, S. R. 1996. North American trilobite species diversity during the Lower Paleozoic. Sixth North American Paleontological Convention Abstracts of Papers. Paleontological Society Special Publication, 8:2.
(4) Adrain, J. M.*, and Edgecombe, G. D. 1994. Silurian encrinurine trilobites of northern Laurentia: systematics of the Struszia genus group. Geological Society of America Abstracts with Programs, 26(7):A373.
(3P) Adrain, J. M.*, and Edgecombe, G. D. 1993. New genera of encrinurid trilobites from the Silurian (Ludlow) Douro Formation of Arctic Canada. Geological Society of America Abstracts with Programs, 25(7):A54.
(2P) Adrain, J. M.* 1992. Silicified Silurian trilobites from the central Canadian Arctic: the most diverse Wenlockian trilobite faunas yet found. Geological Society of America Abstracts with Programs, 24(7):A98.
(1) Adrain, J. M.*, and Chatterton, B. D. E. 1991. Heterochrony in the phylogenesis of otarionine trilobites. Geological Society of America Abstracts with Programs, 23(7):A278.
6. Invited Lectures at Other Institutions
September 2004, University of New Brunswick, Department of Biology, "Community ecology and the fossil record: Can we access ecological information in deep time?"
April 2004, University of Chicago, Department of the Geophysical Sciences, "Paleobiological phenomena and physical context: Microkarst and species turnover in the earliest Ordovician (Skullrockian) of the Great Basin"
April 2002, Carnegie Museum of Natural History, "Early Paleozoic trilobite diversity: phylogenetics, sampling, and global change"
October 2001, Iowa State University, Department of Geological and Atmospheric Sciences, "Early Ordovician trilobite diversity, turnover, and mass extinction"
March, 2001, Keynote Address, Mid-America Paleontological Society National Expo
October, 2000, Harvard University, Department of Earth and Planetary Sciences, "Trilobites, Diversity, and Early Paleozoic Global Change"
7. Pending Decisions
(1) Adrain, J. M.*, Chatterton, B. D. E., and Kloc, G. J. In review. Systematics of the koneprusiine trilobites, with new taxa from the Silurian and Devonian of Laurentia. Journal of Paleontology (submitted August 01, 2005).
TEACHING AT THE UNIVERSITY OF IOWA
|
Semester and
Year |
Course Numbers and Titles
(xx%) indicates
proportion of semester that team course was taught * Indicates
team taught course where I participated in 100% of course |
En-roll-ment |
ACE Student
evaluations of xxx (median score)
Rating Scale: 1
strongly disagree |
# Grad. Advi-sees |
||||
|
Overall
excellent course |
Instruc-tor is
effective |
Course material
present.enthusiastically |
Instructor clearly communicated
material |
|||||
|
Spring 2005 |
012:004/006 |
4/3 |
205 |
5.71 |
5.80 |
5.90 |
5.75 |
4 |
|
Fall 2004 |
012:225 |
1-3 |
13 |
5.95 |
5.96 |
5.85 |
5.83 |
4 |
|
Spring 2004 |
012:004/006 |
4/3 |
208 |
5.65 |
5.84 |
5.94 |
5.70 |
3 |
|
Fall 2003 |
012:144 |
3 |
8 |
6.00 |
6.00 |
5.80 |
5.92 |
3 |
|
|
012:225 |
1-3 |
15 |
5.72 |
5.63 |
5.80 |
5.63 |
|
|
Spring2003 |
012:004 |
4 |
122 |
5.80 |
5.87 |
5.95 |
5.82 |
1 |
|
|
012:006 |
3 |
100 |
5.63 |
5.80 |
5.80 |
5.80 |
|
|
Fall 2002 |
012:225 |
1-3 |
11 |
5.79 |
5.81 |
5.58 |
5.50 |
1 |
|
Spring2002 |
012:006 |
4 |
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